Sergey S. Titov & Anton V. Volynkin
Entomological Studies (ISSN 3153-743X)
Issue: 2 (2026)
Page range: 74–80
DOI: https://doi.org/10.5281/zenodo.24158181
Published: 12 November 2026
Authors' Metadata
Sergey V. Titov
Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ-050060, Almaty, Kazakhstan; &
Pavlodar University named after A. Margulan, Institute of Archaeological Research, 60 Olzhabay Batyr Str., Pavlodar, 140002, Kazakhstan;
https://orcid.org/0000-0003-2196-280X
Anton V. Volynkin
African Natural History Research Trust, Street Court, Leominster, HR6 9QA, United Kingdom; https://orcid.org/0000-0001-9447-4925
Abstract
The paper describes a new species of the Bryophilinae genus Scythobrya Boursin, 1960, Scythobrya kozybayevi Volynkin & Titov, sp. n. from Arkaly Mountains in East Kazakhstan. The diagnostic comparison is made with Scythobrya kaszabi (Pekarsky, Volynkin & Matov, 2014) and Scythobrya miltophaea (Hampson, 1908). Adults and male and female genitalia of the new and similar species are illustrated.
Key words
Arkaly Mountains, Bryophila, Manash Kabashevich Kozybayev, Saur–Tarbagatai Mountains.
How to cite:
Titov, S.V. & Volynkin, A.V. (2026) Scythobrya kozybayevi, a new species from East Kazakhstan (Lepidoptera: Noctuidae: Bryophilinae). Entomological Studies, 2, 74–80.
ES_002_074-080_Titov & Volynkin.pdfReferences
Boursin, Ch. (1954) Zwei neue Cryphia Hb. (Bryophila)-Arten aus dem vorderasiatisch-mediterranen Faunenkreis (Beitrage zur Kenntnis der 'Agrotidae-Trifinae', LXV (66)). Zeitschrift der Wiener Entomologische Gesellschaft, 39, 85–89, 2 pls. [In German]
Hampson, G.F. (1908) Catalogue of the Lepidoptera Phalaenae in the British Museum, 7. Taylor and Francis, London, 709 pp., pls 108–122.
Kononenko, V.S. (2010) Micronoctuidae, Noctuidae: Rivulinae–Agaristinae (Lepidoptera). Noctuidae Sibiricae. Vol. 2. Entomological Press, Sorø, 475 pp.
Lafontaine, J.D. & Mikkola, K. (1987) Las-och-nyckel systemen i de inre genitalierna av Noctuidae (Lepidoptera) som taksonomiska kaennetecken. [Lock-and-key systems in the inner genitalia of Noctuidae (Lepidoptera) as a taxonomic character.] Entomologiske Meddelelser, 55, 161–167. [In Swedish with English summary]
Pekarsky, O., Volynkin, A.V. & Matov, A.Y. (2014) A new species of Bryophila Treitschke, 1825 from Mongolia (Lepidoptera, Noctuidae, Bryophilinae). Zootaxa, 3856 (1), 143–148. http://dx.doi.org/10.11646/zootaxa.3856.1.7
Staudinger, O. (1881) Beitrag zur Lepidopterenfauna Central-Asiens. Stettiner entomologische Zeitung, 42 (10–12), 393–424. [In German]
Volynkin, A.V. (2024) On the terminology of the genitalia structures of lichen moths (Lepidoptera: Erebidae: Arctiinae: Lithosiini) with some references to Noctuidae. Ecologica Montenegrina, 73, 176–207. https://doi.org/10.37828/em.2024.73.18
Author contributions
Sergey V. Titov: Conceptualization, Investigation, Writing—Original draft, Data Curation, Supervision.
Anton V. Volynkin: Investigation, Writing—Original draft, Writing—Review and Editing, Data Curation, Visualization.
Data availability
All of the data that support the study are available in the main text.
Corresponding Author's statements
In submitting this paper (Article), the Corresponding Author confirmed that:
● Corresponding Author has been authorized by the Co-Author, who agreed to submission of the paper.
● The Article has been approved by the Co-Author and by the responsible authorities – tacitly or explicitly – of the institutes where the work has been
carried out.
● This Article has not been published before and is not concurrently being considered for publication elsewhere.
● This Article is not based on any work retracted earlier from any other journal.
Copyright: © Sergey S. Titov & Anton V. Volynkin
This is an open-access article distributed under terms of the Creative Commons Attribution License (Attribution 4.0 International – CC BY 4.0).
In publishing this paper, Authors accept all terms of copyright.
Authors retain copyright and grant the journal right of first publication. Entomological Studies allows others to share the work with an acknowledgement of the work's authorship and initial publication in the journal.
Authors are able to enter into separate, additional contractual arrangements for the non-exclusive distribution of the journal's published version of the work (e.g., post it to an institutional repository or publish it in a book), with an acknowledgement of its initial publication in this journal.
Legal statement
In publishing this paper, Authors confirm that:
● This Article does not violate any copyright or other personal proprietary right of any person or entity and it contains no abusive, defamatory, obscene or fraudulent statements, nor any other statements that are unlawful in any way.
● Authors declare that, to the best of their knowledge, the manuscript complies with the legal requirements of the countries, in which the research was conducted.
Conflict of interest
Authors declare that, to the best of their knowledge, they have no conflict of interest, financial or otherwise, or competing interests.
Artificial Intelligence (AI) use
Authors are accountable and responsible for the contents of their manuscript, including the disclosure of any use of AI.
Authors state that no AI tools were used during preparation of the present paper.
